Family Trochilidae Hummingbirds

Frilled Coquette (Lophornis magnificus) seen in Santa Teresa, Brazil at Pousada Vita Verde, is one of my favorite hummers.

Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 366 species (of which I have seen and documented only 51) and 113 genera, they occur from Alaska to Tierra del Fuego, but most species are found in Central and South America. About 28 hummingbird species are listed as endangered or critically endangered, with numerous species declining in population.

Hummingbirds have varied specialized characteristics to enable rapid, maneuverable flight, exceptional metabolic capacity, adaptations to high altitude, sensitive visual and communication abilities, and long-distance migration in some species. Among all birds, male hummingbirds have the widest diversity of plumage color, particularly in blues, greens, and purples. Hummingbirds are the smallest mature birds, measuring 7.5–13 cm (3–5 in) in length. The smallest is the 5 cm (2.0 in) bee hummingbird, which weighs less than 2.0 g (0.07 oz), and the largest is the 23 cm (9.1 in) giant hummingbird, weighing 18–24 grams (0.63–0.85 oz). Noted for long beaks, hummingbirds are specialized for feeding on flower nectar, but all species also consume small insects.

They are known as hummingbirds because of the humming sound created by their beating wings, which flap at high frequencies audible to other birds and humans. They hover at rapid wing-flapping rates, which vary from around 12 beats per second in the largest species to 80 per second in small hummingbirds.

Hummingbirds have the highest mass-specific metabolic rate of any homeothermic animal. To conserve energy when food is scarce and at night when not foraging, they can enter torpor, a state similar to hibernation, and slow their metabolic rate to 1/15 of its normal rate. While most hummingbirds do not migrate, the rufous hummingbird has one of the longest migrations among birds, traveling twice per year between Alaska and Mexico, a distance of about 3,900 miles (6,300 km).

Little Anna’s Hummingbird (Calypte anna) was the species I grew up with in Southern California where they are still relatively easy to see.

Hummingbirds are restricted to the Americas from south central Alaska to Tierra del Fuego, including the Caribbean. The majority of species occur in tropical and subtropical Central and South America, but several species also breed in temperate climates and some hillstars occur even in alpine Andean highlands at altitudes up to 5,200 m (17,100 ft).

The greatest species richness is in humid tropical and subtropical forests of the northern Andes and adjacent foothills, but the number of species found in the Atlantic Forest, Central America or southern Mexico also far exceeds the number found in southern South America, the Caribbean islands, the United States, and Canada. While fewer than 25 different species of hummingbirds have been recorded from the United States and fewer than 10 from Canada and Chile each, Colombia alone has more than 160 and the comparably small Ecuador has about 130 species.

Hummingbirds are specialized nectarivores tied to the ornithophilous flowers upon which they feed. This coevolution implies that morphological traits of hummingbirds, such as bill length, bill curvature, and body mass, are correlated with morphological traits of plants, such as corolla length, curvature, and volume. Some species, especially those with unusual bill shapes, such as the sword-billed hummingbird and the sicklebills, are coevolved with a small number of flower species. Even in the most specialized hummingbird–plant mutualisms, the number of food plant lineages of the individual hummingbird species increases with time. The bee hummingbird (Mellisuga helenae) – the world's smallest bird – evolved to dwarfism likely because it had to compete with long-billed hummingbirds having an advantage for nectar foraging from specialized flowers, consequently leading the bee hummingbird to more successfully compete for flower foraging against insects.

Long-tailed Sylph (Aglaiocercus kingii) is one of the more spectacular species in the Ecuadorian Elfin forests. Pretty common too!

Many plants pollinated by hummingbirds produce flowers in shades of red, orange, and bright pink, although the birds take nectar from flowers of other colors. Hummingbirds can see wavelengths into the near-ultraviolet, but hummingbird-pollinated flowers do not reflect these wavelengths as many insect-pollinated flowers do. This narrow color spectrum may render hummingbird-pollinated flowers relatively inconspicuous to most insects, thereby reducing nectar robbing. Hummingbird-pollinated flowers also produce relatively weak nectar (averaging 25% sugars) containing a high proportion of sucrose, whereas insect-pollinated flowers typically produce more concentrated nectars dominated by fructose and glucose.

Hummingbirds and the plants they visit for nectar have a tight coevolutionary association, generally called a plant–bird mutualistic network. These birds show high specialization and modularity, especially in communities with high species richness. These associations are also observed when closely related hummingbirds, such as two species of the same genus, visit distinct sets of flowering species.

The galleries below have many images of thee 51 species I have been fortunate to have documented over the years. I suspect that I have seen at least that many again while traveling in Panama, Mexico, Costa Rica, Belize, Guatemala, Nicaragua & Columbia before I began keeping lists or even bothering to snap a picture.

Identification in this group is difficult for me, so many of the ID’s need to be considered to be tentative. Please feel free to correct me if you see any errors - much appreciated. Cheers